Pace in races vs pace in training

Something has been bugging me for a while, and thats the perception of effort at a given pace in training vs the same, or similar pace in races, and the differences, is it a purely physical thing, a mental thing, or a combination of the two.?

Last night I did a 5 mile speed session, this session called for 2 reps of 1 miles @ 9:10 pace, towards the end of the 2nd rep, I was REALLY struggling, it was really hard, and I felt like I was pushing almost as hard as I could.

Interestingly, mile 5 to mile 6 of my last 10k race was paced at almost exactly the same pace, however, I can't remember that mile feeling anywhere near as tough as the mile last night.

Why is that?

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